Funding for this research was provided by:
John Templeton Foundation (54281)
National Science Foundation
NSF Graduate Research Fellowship
Received: 26 January 2017
Accepted: 26 July 2017
First Online: 24 August 2017
Ethics approval and consent to participate
: All field procedures were approved by the University of Utah Institutional Animal Care and Use Committee (protocols #07–08004, #10–07003 and #13–06010) and by the Galápagos National Park.
: The authors declare that they have no competing interests.
: Springer Nature remains neutral with regard to jurisdictional claims in published maps and institutional affiliations.